(These together sometimes are called the lower
deuterostomes.[3])
Whether the
Xenacoelomorpha clade is the sister group to the Ambulacraria remains a contentious issue, with some authors arguing that the former should be placed more basally among
metazoans,[4][5][6] and other authors asserting that the best choices of phylogenetic methods support the position of Xenacoelomorpha as the sister group to Ambulacraria.[7][8]
As for many animals, the
egg cell of any
extant ambulacrarian divides and develops into a
blastula ("cell ball"), which develops into a
triploblast ("three-layered")
gastrula. The gastrula then develops into a
dipleurula larva form in the Asteroidea, Holothuroidea, Crinoidea, and Hemichordata, and into a
pluteus larva form in the Echinoidea and Ophiuroidea.[3][10] This, in its turn, is developed in various different kinds of larvae for different taxa of ambulacrarians.
It has been suggested that the adult form of the
last common ancestor of the ambulacrarians was
anatomically similar to the dipleurula larva; this hypothetic ancestor sometimes also is called dipleurula.[11]
^
abLacalli, Thurston Castle.
"Tutorial". Marine Invertebrate larvae: A study in morphological diversity.
University of Saskatchewan. Retrieved 2020-01-13.