An approximate timescale of the Cenozoic.
Axis scale: millions of years ago.
The European Land Mammal Mega Zones (abbreviation: ELMMZ, more commonly known as European land mammal ages or ELMA) are zones in rock layers that have a specific assemblage of fossils (
biozones) based on occurrences of fossil assemblages of
European land
mammals. These biozones cover most of the
Cenozoic, with particular focus having been paid to the
Neogene and
Paleogenesystems (i.e. rock layers which are 65.5 to 2.588 million years old), the
Quaternary has several competing systems. In cases when fossils of mammals are abundant,
stratigraphers and
paleontologists can use these biozones as a more practical regional alternative to the
stages of the official
ICSgeologic timescale. European Land Mammal Mega Zones are often also confusingly referred to as ages, stages, or intervals.[1]
Biostratigraphic methods
Mammal zones were, like all biozones, established using geographic place names where
fossil materials were obtained. The basic unit of measure is the first/last boundary statement. This shows that the first appearance event of one
taxon is known to predate the last appearance event of another. If two
taxa are found in the same fossil
quarry or at the same
stratigraphic horizon, then their age-range zones overlap.
The terrestrial stratigraphy of the
Cenozoic is more difficult than that of marine deposits. The geologic timescale of the ICS is therefore based on marine fossils, that don't occur in terrestrial sediments. This makes the
correlation of terrestrial deposits with the ICS timescale often difficult. Correlation is possible when marine deposits interfinger with terrestrial deposits (resulting from a series of
transgressions and
regressions of the sea during deposition), but this isn't the case everywhere. A fine stratigraphic division of the terrestrial record can in most places only be made using fossils of land species. Small mammals are often the best choice as they are quite abundant in the terrestrial record, especially their teeth. Teeth have an even better chance of preservation than bones.
The European mammalian biozones were established for the
Paleogene (66-23.03
Mya, 8 zones) and
Neogene (23.03-2.58
Mya, 7 zones) separately. Some of these, especially for the Neogene, were already established in the 19th century. The
Villafranchian was, for example, introduced by
Lorenzo Pareto in 1865. A finer subdivision was established by
Pierre Mein in 1975, who divided the Neogene in 17 zones, known as the
MN zonation, indicated by the letters MN (Mammal Neogene) and a number.
Similarly, a more detailed subdivision for the Paleogene period was established. There are 30 such
Mammal Paleogene zones (MP1 to MP30, numbered from old to young).[2]
European Land Mammal Mega Zones most often have their bases at first appearances (FAD, First Appearance Date) of a certain
species or
genus. The numbers are higher for younger zones. Due to a redefinition of the boundary between the Neogene and
Quaternary periods, MN 17 is now in fact considered a Quaternary biozone.
The first zonation for the
Quaternary of Europe was proposed by Azzaroli in 1967.[3] This was then expanded by Gliozzi et al. in 1997 to make a system of 3 'ages' subdivided into 13 'faunal units'.[4] The scheme does not define boundaries but instead is accompanied by a range chart, where the entry and exit dates for the taxa are indicated. Each zone is named after a reference locality. Most of the reference locations are in Italy but the scheme is used in other European regions. The mammal ages and Faunal Units (FU) after Gliozzi et al. are:
In 1982, Guérin[10] proposed an alternative scheme, which extended the
MN zonation scheme for the
Neogene with additional units to cover the Quaternary. There have been further updates since.[11] The MNQ (Mammal Neogene Quaternary) scheme added an additional 12 units in total, MNQ 16–27.
For small mammals there is a third scheme, the MmQ, published by Agustí, Moyà‐Solà, and Pons‐Moyà in 1987.[12] The scheme includes some large mammals for reference and thus has a wider application.[13]
^Azzaroli, Augusto. 1967. “Villafranchian Correlations Based on Large Mammals.” Giornale di Geologia 35 (1): 21
^
abGliozzi, Elsa, Laura Abbazzi, Patrizia Argenti, Augusto Azzaroli, Lucia Caloi, Lucia Capasso Barbato, Giuseppe Di Stefano, et al. 1997. “Biochronology of Selected Mammals, Molluscs and Ostracods from the Middle Pliocene to the Late Pleistocene in Italy: The State of the Art.” Rivista Italiana di Paleontologia e Stratigrafia 103 (3): 369–88.
^Kurten, Bjorn (1968). Pleistocene Mammals of Europe. London: Weidenfeld and Nicolson.
^Croitor, Roman (2018). Plio-Pleistocene Deer of Western Palearctic: Taxonomy, Systematics, Phylogeny. Institute of Zoology of the Academy of Sciences of Moldova.
ISBN978-9975-66-609-1.
OCLC1057238213.
^Guérin, Claude. 1982. “Première Biozonation du Pléistocène Européen, Principal Résultat Biostratigraphique de l'Étude des Rhinocerotidae (Mammalia, Perissodactyla) du Miocène Terminal au Pléistocène Supérieur d'Europe Occidentale.” Geobios 15 (4): 593–98. DOI:10.1016/s0016-6995(82)80076-4
^Faure, Martine, and Claude Guérin. 1992. “La Grande Faune d'Europe Occidentale au Pléistocène Moyen et Supérieur et ses Potentialités d'Information en Préhistoire.” Mémoires de la Société Géologíque de France 160: 77–84.
^Agustí, Jordi, Salvador Moyà‐Solà, and Joan Pons‐Moyà. 1987. “La Sucesión de Mamíferos en el Pleistoceno Inferior de Europa: Proposición de Una Nueva Escala Bioestratigráfica.” Paleontología i Eovlució, mem. esp. 1: 287–95.
^van der, Made, J. (2018). Quaternary Large‐Mammal Zones. In The Encyclopedia of Archaeological Sciences, S.L. López Varela (Ed.). doi:10.1002/9781119188230.saseas0486
Notes
Literature
Koufos, G.D.; Kostopoulos, D.S. & Vlachou, T.D. (2005). '"Neogene/Quaternary mammalian migrations in Eastern Mediterranean", Belgian Journal of Zoology135(2): pp. 181–190.
Mein, P. (1975). Report on activity RCMNS-Working groups, 1971–1975, pp. 78–81, Bratislava.
Rose, Kenneth D. (2006). The beginning of the age of mammals. Baltimore, Md.: Johns Hopkins Univ. Press.
ISBN978-0801884726.
Steininger, F.F. (1999). "Chronostratigraphy, Geochronology and Biochronology of the Miocene "European Land Mammal Mega-Zones" (ELMMZ) and the Miocene "Mammal Zones (MN-Zones)"". In Rössner, G. E.; Heissig, K. (eds.). The Miocene land mammals of Europe. München: Pfeil. pp. 9–24.
ISBN3-931516-50-4.