Pervushovisaurus Temporal range:
Cenomanian
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P. campylodon snout fragment, the lectotype specimen of the species | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Ichthyosauria |
Family: | † Ophthalmosauridae |
Subfamily: | † Platypterygiinae |
Genus: | †
Pervushovisaurus Arkhangel'skii, 1998 |
Species | |
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Synonyms | |
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Pervushovisaurus is a genus of platypterygiine ichthyosaur from the Late Cretaceous ( Cenomanian) of the Saratov region in western Russia, the La Penthiève Beds of France and the Cambridge area of the UK.
It was originally described as a subgenus of Platypterygius, [1] but later work showed that it was sufficiently distinct from the type species of Platypterygius, P. platydactylus, to be elevated to full generic rank. [2] The type species of Pervushovisaurus, P. bannovkensis, is known only from the holotype, SSU 104a/24, a partial skull. [3] In 2016, an additional species of Platypterygius from England and France, P. campylodon, was also referred to Pervushovisaurus after previously being allied with Ichthyosaurus in 1846. [4] [5] [6] A previous Russian species of Platypterygius, "P." kiprianoffi, was also assigned to P. campylodon. [7] [8] [9]
A specimen of P. campylodon was found in the La Penthiève Beds of France and was described in 1882 by Henri Émile Sauvage. [10]
Pervushovisaurus was a medium-sized ichthyosaur measuring 5.8 metres (19 ft) long. [1] Fishcer (2016) gave the emended diagnosis of Pervushovisaurus as follows, based on that of Fischer et al. (2014): [2] [6] "Platypterygiine ophthalmosaurid characterized by the following autapomorphies [...]: presence of foramina along the ventral premaxillary–maxillary suture; presence of a semi-oval foramen on the lateral surface of the premaxilla, anteroventral to the external naris; presence of lateral ridges on the maxilla; presence of wide supranarial ‘wing’ of the nasal (a similar structure, although much smaller, is present in ‘Platypterygius’ australis and Acamptonectes densus) (see Kear, 2005; Fischer et al., 2012, respectively); robust splenial markedly protruding from the external surface of the mandible; root with quadrangular cross-section, with the cementum forming prominent 90° angles.
Pervushovisaurus is also characterized by the following unique combination of features: secondarily closed naris surrounded by foramina (as in ‘Platypterygius’ sachicarum and ‘Platypterygius’ australis (see Paramo, 1997; Kear, 2005, respectively), and in Simbirskiasaurus birjukovi, although the ‘anterior’ naris is still present in this taxon (Maisch & Matzke, 2000; Fischer et al., 2014a)); elongated anterior process of the maxilla, reaching anteriorly the level of the nasal (unlike in Aegirosaurus leptospondylus, Sveltonectes insolitus and Muiscasaurus catheti) (Bardet & Fernández, 2000; Fischer et al., 2011a; Maxwell et al., 2015, respectively); rostrum straight (unlike in ‘Platypterygius’ americanus, ‘Platypterygius’ sachicarum, ‘Platypterygius’ australis and possibly Muiscasaurus catheti, where it is slightly curved anteroventrally Romer, 1968; Paramo, 1997; Kear, 2005; Maxwell et al., 2015, respectively); straight, non-recurved tooth crowns (unlike in Sveltonectes insolitus, Muiscasaurus catheti) (Fischer et al., 2011a; Maxwell et al., 2015, respectively)."