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Jenny Clack

Born
Jennifer Alice Agnew

(1947-11-03)3 November 1947
Died26 March 2020(2020-03-26) (aged 72)
NationalityEnglish
Alma mater Newcastle University (BSc, PhD)
University of Leicester
University of Cambridge (MA)
Known forGaining Ground: the Origin and Early Evolution of Tetrapods (2002)
Scientific career
Fields Zoology
Palaeontology
Evolutionary biology
Institutions

Jennifer Alice Clack, FRS, FLS (née Agnew; 3 November 1947 – 26 March 2020) was an English palaeontologist and evolutionary biologist. She specialised in the early evolution of tetrapods, specifically studying the "fish to tetrapod" transition: the origin, evolutionary development and radiation of early tetrapods and their relatives among the lobe-finned fishes. She is best known for her book Gaining Ground: the Origin and Early Evolution of Tetrapods, published in 2002 (second edition, 2012) and written with the layperson in mind.

Clack was curator at the Museum of Zoology and Professor of Vertebrate Palaeontology at the University of Cambridge, where she devoted her career to studying the early development of tetrapods, the "four-legged" animals said to have evolved from Devonian lobe-finned fishes and colonised the freshwater swamps of the Carboniferous period.

Early life and education

Clack was born on 3 November 1947, the only child of Ernest and Alice Agnew. [1] She was brought up in Manchester, England. [2] She was educated at Bolton School (Girls' Division), a private school in Bolton, Lancashire. She received a B.Sc. in Zoology from the University of Newcastle upon Tyne in 1970. In 1978, she accepted an invitation from Alec Panchen to obtain a Ph.D. from the same university, at the encouragement of her then new boyfriend Rob Clack. They married in 1980. Her doctorate was completed in 1984. She also held a Graduate Certificate in Museum Studies from the University of Leicester and an MA from the University of Cambridge. On 9 December 2000, she was awarded a Doctor of Science (ScD) degree by the University of Cambridge. [3]

Academic career

In 1981, Clack joined the University Museum of Zoology, University of Cambridge, as an Assistant Curator. [1] [2] She was promoted to Senior Assistant Curator in 1995. Since 2005, she has been Curator in Vertebrate Palaeontology at the museum. [1] In 2006, she was awarded a personal chair by the University of Cambridge, and took the title Professor of Vertebrate Palaeontology. [4] She retired in 2015 and became Emeritus Professor of Vertebrate Palaeontology at the museum. [5] In 1997, Clack was elected a Fellow of Darwin College, Cambridge; [6] since 1 October 2015, she has been an Emeritus Fellow. [7] [8] From 2000 to 2005, she was Reader in Vertebrate Palaeontology at the University of Cambridge. [1] On 1 October 2006, Clack was awarded a personal chair by Cambridge, taking the title Professor of Vertebrate Palaeontology. [1] [9]

Clack is best known for her extensive body of work on early tetrapods, much of which redefined how paleontologists conceived of the evolution of limbs and other features associated with tetrapods' transition from other lobe-finned fishes. She began her career working primarily on the ear of early tetrapods [10] and later expanded to more broadly addressing the osteology and evolution of tetrapods. Together with Michael Coates ( University of Chicago), Clack defined what is known as " Romer's Gap", [11] a major gap in the fossil record of early tetrapods and one that she subsequently began to fill in collaboration with other paleontologists. Clack also undertook extensive fieldwork expeditions in order to search for further remains of early tetrapods. In 1987, during an expedition to East Greenland, Clack and her team discovered the remains of the Devonian tetrapods Acanthostega and Ichthyostega, following up on field notes of researchers who had collected material of Acanthostega in 1970. [12] Additional surveys in 1998 led to the collection of substantial new material, including what is now recognized as Ymeria. [13] Most recently, she led a major consortium project (TW:eed [14]) investigating some exciting new fossils from Northumberland and the Borders Region of Scotland which date from the Tournaisian stage of the earliest Carboniferous period; this project has produced numerous publications furthering the understanding of early tetrapod evolution. [15] [16] [17] [18]

Over the course of her lengthy career, Clack published in some of the most notable scientific journals, including Nature, [19] [20] [21] [22] [23] [24] [25] [26] [27] [28] [29] [30] [31] [32] [33] Science, [34] [35] and Proceedings of the National Academy of Sciences [36] [37] and is one of the most published vertebrate paleontologists in Nature, arguably the leading scientific journal in the world, with over 15 papers in that journal alone. In addition to her Gaining Ground book, Clack also co-authored a volume of the Handbuch der Paläoherpetologie series on early tetrapods with Andrew Milner in 2015 [38] and co-edited a volume on the evolution of hearing in 2016. [39]

Clack is probably best known for discovering that the earliest tetrapods had more than five toes per foot: the Upper Devonian tetrapods from East Greenland, Ichthyostega had seven while Acanthostega had eight (compared to the six toes of the Russian Devonian tetrapod Tulerpeton). This suggests that pentadactyly was not an ancestral trait for tetrapods. [40]

Clack supervised many graduate students who went on to pursue successful careers in paleontology and evolutionary biology, including Per Ahlberg ( Uppsala University), Paul Upchurch ( University College London), Michael Lee ( Flinders University), and Matthew Friedman ( University of Michigan). [5] In April 2012 she was featured in an episode [41] of the BBC television series Beautiful Minds, a set of documentaries about scientists who have made important discoveries. This may be viewed on YouTube. [42] Clack was honored by her peers with a festschrift published in 2019. [5]

Taxa named by Jennifer Clack and colleagues
Year Taxon Authors
2020 Rossichthys clackae gen. et sp. nov Johanson et al. [43]
2020 Brittagnathus minutus gen. et sp. nov. Ahlberg & Clack [44]
2019 Parmastega aelidae gen. et sp. nov. Beznosov, Clack, Lukševičs, Ruta & Ahlberg [33]
2019 Limanichthys fraseri gen. et sp. nov. Challands et al. [45]
2018 Celsiodon ahlbergi gen. et sp. nov. Clack, Challands, Smithon & Smithson [46]
2018 Mesanerpeton woodi gen. et sp. nov. Smithson & Clack [47]
2018 Whitropus longicalcus gen. et sp. nov. Richards et al. [48]
2018 Deltodus tubineus sp. nov. Richards et al. [48]
2017 Spathicephalus marsdeni sp. nov. Smithson et al. [49]
2016 Perittodus apsconditus gen. et sp. nov. Clack & Smithson [50]
2016 Koilops herma gen. et sp. nov. Clack & Smithson [50]
2016 Ossirarus kierani gen. et sp. nov. Clack & Smithson [50]
2016 Diploradus austiumensis gen. et sp. nov. Clack & Smithson [50]
2016 Aytonerpeton microps gen. et sp. nov. Clack & Smithson [50]
2015 Ctenodus williei sp. nov. Smithson, Richards & Clack [51]
2015 Ctenodus whitropei sp. nov. Smithson, Richards & Clack [51]
2015 Ctenodus roberti sp. nov. Smithson, Richards & Clack [51]
2015 Xylognathus macrustenus gen. et sp. nov. Smithson, Richards & Clack [51]
2015 Ballagadus rossi gen. et sp. nov. Smithson, Richards & Clack [51]
2015 Ballagadus caustrimi sp. nov. Smithson, Richards & Clack [51]
2015 Coccovedus celatus gen. et sp. nov. Smithson, Richards & Clack [51]
2015 Occludus romeri gen. nov. Smithson, Richards & Clack [51]
2012 Ymeria denticulata gen. et sp. nov. Clack, Ahlberg, Blöm & Finney [52]
2011 Kirktonecta milnerae gen. et sp. nov. Clack [53]
2004 Occidens portlocki gen. et sp. nov. Clack & Ahlberg [54]
2003 Kyrinion martilli gen. et sp. nov. Clack [55]
2002 Pederpes finneyae gen. et sp. nov. Clack [23]
1998 Eucritta melanolimnetes gen. et sp. nov. Clack [22]
1993 Silvanerpeton miripedes gen. et sp. nov. Clack [56]

Death

Clack died on 26 March 2020 at the age of 72, after a five-year battle with endometrial cancer. [57]

Honours

In 2008, Clack was awarded the Daniel Giraud Elliot Medal from the U.S. National Academy of Sciences, the first woman to achieve the honor. [58]

In 2009, Clack was elected a Fellow of the Royal Society, [59] the first female vertebrate paleontologist to achieve the honor. [60] She has also been elected a Foreign Honorary Member of the American Academy of Arts and Sciences. [61]

On 15 June 2013, Clack was awarded an honorary Doctor of Science (DSc) degree by the University of Chicago. The university described her as "an internationally preeminent palaeontologist whose research has profoundly changed the understanding of the origin of terrestrial vertebrate life." [62] Also in 2013, she was awarded the T Neville George Medal by the Geological Society of Glasgow. [63]

On 17 July 2014, she was awarded an honorary Doctor of Science degree by the University of Leicester. [64] Also in 2014, she was made an Honorary Foreign Member of the Royal Swedish Academy of Sciences. [65]

In 2018, she won the Palaeontological Association's most prestigious award, the Lapworth Medal. [66]

References

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  2. ^ a b Clack, Rob (6 October 2009). "Jenny Clack – Biography". The Clacks. Retrieved 3 November 2015.
  3. ^ "Congregation of the Regent House on 9 December 2000". Cambridge University Reporter. No. 5831. 13 December 2000. p. 27. Retrieved 3 November 2015.
  4. ^ "Clack, Prof. Jennifer Alice, (born 3 Nov. 1947), Curator in Vertebrate Palaeontology, 2005–15, and Professor, 2006–15, now Professor Emeritus of Vertebrate Palaeontology, University Museum of Zoology, Cambridge; Fellow, Darwin College, Cambridge, 1997–2015, now Emeritus". Who Was Who. Oxford University Press. 1 December 2020. doi: 10.1093/ww/9780199540884.013.U249891.
  5. ^ a b c Ruta, Marcello; Ahlberg, Per E.; Smithson, Timothy R. (2018). "Fossils, function and phylogeny: Papers on early vertebrate evolution in honour of Professor Jennifer A. Clack – Introduction". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 109 (1–2): 1–14. doi: 10.1017/S1755691019000057. ISSN  1755-6910.
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  10. ^ Clack, J. A. (1983). "The stapes of the Coal Measures embolomere Pholiderpeton scutigerum Huxley (Amphibia: Anthracosauria) and otic evolution in early tetrapods". Zoological Journal of the Linnean Society. 79 (2): 121–148. doi: 10.1111/j.1096-3642.1983.tb01163.x. ISSN  0024-4082.
  11. ^ Coates, Michael I.; Clack, Jennifer A. (1995). "Romer's gap: Tetrapod origins and terrestriality". Bulletin du Muséum National d'Histoire Naturelle, Section C. 4e série. 17: 373–388.
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  13. ^ CLACK, JENNIFER A.; AHLBERG, PER E.; BLOM, HENNING; FINNEY, SARAH M. (2012). "A new genus of Devonian tetrapod from North-East Greenland, with new information on the lower jaw of Ichthyostega". Palaeontology. 55 (1): 73–86. Bibcode: 2012Palgy..55...73C. doi: 10.1111/j.1475-4983.2011.01117.x. ISSN  0031-0239.
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  15. ^ Clack, Jennifer A.; Bennett, Carys E.; Carpenter, David K.; Davies, Sarah J.; Fraser, Nicholas C.; Kearsey, Timothy I.; Marshall, John E. A.; Millward, David; Otoo, Benjamin K. A.; Reeves, Emma J.; Ross, Andrew J. (5 December 2016). "Phylogenetic and environmental context of a Tournaisian tetrapod fauna" (PDF). Nature Ecology & Evolution. 1 (1): 2. doi: 10.1038/s41559-016-0002. ISSN  2397-334X. PMID  28812555. S2CID  22421017.
  16. ^ Otoo, Benjamin K. A.; Clack, Jennifer A.; Smithson, Timothy R.; Bennett, Carys E.; Kearsey, Timothy I.; Coates, Michael I. (3 October 2018). "A fish and tetrapod fauna from Romer's Gap preserved in Scottish Tournaisian floodplain deposits". Palaeontology. 62 (2): 225–253. doi: 10.1111/pala.12395. ISSN  0031-0239.
  17. ^ Smithson, Timothy R.; Clack, Jennifer A. (2017). "A new tetrapod from Romer's Gap reveals an early adaptation for walking". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 108 (1): 89–97. doi: 10.1017/s1755691018000075. ISSN  1755-6910. S2CID  232149117.
  18. ^ Clack, Jennifer A.; Bennett, Carys E.; Davies, Sarah J.; Scott, Andrew C.; Sherwin, Janet E.; Smithson, Timothy R. (3 January 2019). "A Tournaisian (earliest Carboniferous) conglomerate-preserved non-marine faunal assemblage and its environmental and sedimentological context". PeerJ. 6: e5972. doi: 10.7717/peerj.5972. ISSN  2167-8359. PMC  6321757. PMID  30627480.
  19. ^ Clack, J. A. (1989). "Discovery of the earliest-known tetrapod stapes". Nature. 342 (6248): 425–427. Bibcode: 1989Natur.342..425C. doi: 10.1038/342425a0. ISSN  0028-0836. PMID  2586610. S2CID  4348808.
  20. ^ Coates, M. I.; Clack, J. A. (1991). "Fish-like gills and breathing in the earliest known tetrapod". Nature. 352 (6332): 234–236. Bibcode: 1991Natur.352..234C. doi: 10.1038/352234a0. ISSN  0028-0836. S2CID  4340202.
  21. ^ Ahlberg, Per E.; Clack, Jennifer A.; Luks̆evic̆s, Ervīns (1996). "Rapid braincase evolution between Panderichthys and the earliest tetrapods". Nature. 381 (6577): 61–64. Bibcode: 1996Natur.381...61A. doi: 10.1038/381061a0. ISSN  0028-0836. S2CID  4338219.
  22. ^ a b Clack, Jennifer A. (1998). "A new Early Carboniferous tetrapod with a mélange of crown-group characters". Nature. 394 (6688): 66–69. Bibcode: 1998Natur.394...66C. doi: 10.1038/27895. ISSN  0028-0836. S2CID  204998306.
  23. ^ a b Clack, J. A. (2002). "An early tetrapod from 'Romer's Gap'". Nature. 418 (6893): 72–76. Bibcode: 2002Natur.418...72C. doi: 10.1038/nature00824. ISSN  0028-0836. PMID  12097908. S2CID  741732.
  24. ^ Clack, J. A.; Ahlberg, P. E.; Finney, S. M.; Dominguez Alonso, P.; Robinson, J.; Ketcham, R. A. (2003). "A uniquely specialized ear in a very early tetrapod". Nature. 425 (6953): 65–69. Bibcode: 2003Natur.425...65C. doi: 10.1038/nature01904. ISSN  0028-0836. PMID  12955140. S2CID  4411060.
  25. ^ Clément, Gaël; Ahlberg, Per E.; Blieck, Alain; Blom, Henning; Clack, Jennifer A.; Poty, Edouard; Thorez, Jacques; Janvier, Philippe (2004). "Devonian tetrapod from western Europe". Nature. 427 (6973): 412–413. doi: 10.1038/427412a. ISSN  0028-0836. PMID  14749820. S2CID  4414996.
  26. ^ Ahlberg, Per Erik; Clack, Jennifer A.; Blom, Henning (2005). "The axial skeleton of the Devonian tetrapod Ichthyostega". Nature. 437 (7055): 137–140. Bibcode: 2005Natur.437..137A. doi: 10.1038/nature03893. ISSN  0028-0836. PMID  16136143. S2CID  4370488.
  27. ^ Ahlberg, Per Erik; Clack, Jennifer A. (2006). "A firm step from water to land". Nature. 440 (7085): 748–749. doi: 10.1038/440747a. ISSN  0028-0836. PMID  16598240. S2CID  4392361.
  28. ^ Ahlberg, Per E.; Clack, Jennifer A.; Lukševičs, Ervīns; Blom, Henning; Zupiņš, Ivars (2008). "Ventastega curonica and the origin of tetrapod morphology". Nature. 453 (7199): 1199–1204. Bibcode: 2008Natur.453.1199A. doi: 10.1038/nature06991. ISSN  0028-0836. PMID  18580942. S2CID  4344417.
  29. ^ Pierce, Stephanie E.; Clack, Jennifer A.; Hutchinson, John R. (23 May 2012). "Three-dimensional limb joint mobility in the early tetrapod Ichthyostega". Nature. 486 (7404): 523–526. Bibcode: 2012Natur.486..523P. doi: 10.1038/nature11124. ISSN  0028-0836. PMID  22722854. S2CID  3127857.
  30. ^ Pierce, Stephanie E.; Ahlberg, Per E.; Hutchinson, John R.; Molnar, Julia L.; Sanchez, Sophie; Tafforeau, Paul; Clack, Jennifer A. (13 January 2013). "Vertebral architecture in the earliest stem tetrapods" (PDF). Nature. 494 (7436): 226–229. Bibcode: 2013Natur.494..226P. doi: 10.1038/nature11825. ISSN  0028-0836. PMID  23334417. S2CID  4329395.
  31. ^ Clack, Jennifer A. (2015). "The origin of terrestrial hearing". Nature. 519 (7542): 168–169. doi: 10.1038/519168a. ISSN  0028-0836. PMID  25762279. S2CID  4392314.
  32. ^ Sanchez, Sophie; Tafforeau, Paul; Clack, Jennifer A.; Ahlberg, Per E. (2016). "Life history of the stem tetrapod Acanthostega revealed by synchrotron microtomography". Nature. 537 (7620): 408–411. Bibcode: 2016Natur.537..408S. doi: 10.1038/nature19354. ISSN  0028-0836. PMC  6485594. PMID  27602519.
  33. ^ a b Beznosov, Pavel A.; Clack, Jennifer A.; Lukševičs, Ervīns; Ruta, Marcello; Ahlberg, Per Erik (2019). "Morphology of the earliest reconstructable tetrapod Parmastega aelidae" (PDF). Nature. 574 (7779): 527–531. Bibcode: 2019Natur.574..527B. doi: 10.1038/s41586-019-1636-y. ISSN  0028-0836. PMID  31645719. S2CID  204848799.
  34. ^ Clack, J. A. (2 April 2004). "PALEONTOLOGY: Enhanced: From Fins to Fingers". Science. 304 (5667): 57–58. doi: 10.1126/science.1096415. ISSN  0036-8075. PMID  15060312. S2CID  82119710.
  35. ^ Callier, V.; Clack, J. A.; Ahlberg, P. E. (17 April 2009). "Contrasting Developmental Trajectories in the Earliest Known Tetrapod Forelimbs". Science. 324 (5925): 364–367. Bibcode: 2009Sci...324..364C. doi: 10.1126/science.1167542. ISSN  0036-8075. PMID  19372425. S2CID  28461841.
  36. ^ Smithson, T. R.; Wood, S. P.; Marshall, J. E. A.; Clack, J. A. (5 March 2012). "Earliest Carboniferous tetrapod and arthropod faunas from Scotland populate Romer's Gap". Proceedings of the National Academy of Sciences. 109 (12): 4532–4537. Bibcode: 2012PNAS..109.4532S. doi: 10.1073/pnas.1117332109. ISSN  0027-8424. PMC  3311392. PMID  22393016.
  37. ^ Neenan, James M.; Ruta, Marcello; Clack, Jennifer A.; Rayfield, Emily J. (22 April 2014). "Feeding biomechanics in Acanthostega and across the fish–tetrapod transition". Proceedings of the Royal Society B: Biological Sciences. 281 (1781): 20132689. doi: 10.1098/rspb.2013.2689. ISSN  0962-8452. PMC  3953833. PMID  24573844.
  38. ^ "Basal Tetrapoda". Dr. Friedrich Pfeil Publishing. Retrieved 12 July 2020.
  39. ^ Clack, Jennifer A.; Fay, Richard R; Popper, Arthur N., eds. (2016). Evolution of the Vertebrate Ear. Springer Handbook of Auditory Research. Vol. 59. Cham: Springer International Publishing. doi: 10.1007/978-3-319-46661-3. ISBN  978-3-319-46659-0. S2CID  36018536.
  40. ^ Coates, M. I.; Clack, J. A. (1990). "Polydactyly in the earliest known tetrapod limbs". Nature. 347 (6288): 66–69. Bibcode: 1990Natur.347...66C. doi: 10.1038/347066a0. ISSN  0028-0836. S2CID  4319165.
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  43. ^ Johanson, Zerina; Jeffery, Jonathan; Challands, Tom; Pierce, Stephanie; Clack, Jennifer (27 October 2020). "A New Look At Carboniferous Rhizodontid Humeri (Sarcopterygii; Tetrapodomorpha)". Journal of Vertebrate Paleontology. e1813150 (3): e1813150. Bibcode: 2020JVPal..40E3150J. doi: 10.1080/02724634.2020.1813150. S2CID  227241079.
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  45. ^ Challands, Tom J.; Smithson, Timothy R.; Clack, Jennifer A.; Bennett, Carys E.; Marshall, John E. A.; Wallace-Johnson, Sarah M.; Hill, Henrietta (11 March 2019). "A lungfish survivor of the end-Devonian extinction and an Early Carboniferous dipnoan radiation". Journal of Systematic Palaeontology. 17 (21): 1825–1846. doi: 10.1080/14772019.2019.1572234. ISSN  1477-2019. S2CID  91623116.
  46. ^ Clack, Jennifer Alice; Challands, Thomas James; Smithson, Timothy Richard; Smithson, Keturah Zoe (2 November 2018). "Newly recognized Famennian lungfishes from East Greenland reveal tooth plate diversity and blur the Devonian–Carboniferous boundary". Papers in Palaeontology. 5 (2): 261–279. doi: 10.1002/spp2.1242. ISSN  2056-2802. S2CID  134074159.
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  48. ^ a b Richards, Kelly R.; Sherwin, Janet E.; Smithson, Timothy R.; Bennion, Rebecca F.; Davies, Sarah J.; Marshall, John E. A.; Clack, Jennifer A. (2017). "Diverse and durophagous: Early Carboniferous chondrichthyans from the Scottish Borders". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 108 (1): 67–87. doi: 10.1017/s1755691018000166. ISSN  1755-6910. S2CID  133929085.
  49. ^ Smithson, Timothy R.; Browne, Michael A. E.; Davies, Sarah J.; Marshall, John E. A.; Millward, David; Walsh, Stig A.; Clack, Jennifer A. (2017). "A new Mississippian tetrapod from Fife, Scotland, and its environmental context". Papers in Palaeontology. 3 (4): 547–557. doi: 10.1002/spp2.1086. hdl: 2381/40472. ISSN  2056-2802.
  50. ^ a b c d e Clack, Jennifer A.; Bennett, Carys E.; Carpenter, David K.; Davies, Sarah J.; Fraser, Nicholas C.; Kearsey, Timothy I.; Marshall, John E. A.; Millward, David; Otoo, Benjamin K. A.; Reeves, Emma J.; Ross, Andrew J. (5 December 2016). "Phylogenetic and environmental context of a Tournaisian tetrapod fauna" (PDF). Nature Ecology & Evolution. 1 (1): 2. doi: 10.1038/s41559-016-0002. ISSN  2397-334X. PMID  28812555. S2CID  22421017.
  51. ^ a b c d e f g h Smithson, Timothy R.; Richards, Kelly R.; Clack, Jennifer A. (2016). "Lungfish diversity in Romer's Gap: reaction to the end-Devonian extinction". Palaeontology. 59 (1): 29–44. Bibcode: 2016Palgy..59...29S. doi: 10.1111/pala.12203. ISSN  1475-4983.
  52. ^ Clack, Jennifer A.; Ahlberg, Per E.; Blom, Henning; Finney, Sarah M. (2012). "A new genus of Devonian tetrapod from North-East Greenland, with new information on the lower jaw of Ichthyostega: A NEW GENUS OF DEVONIAN TETRAPOD". Palaeontology. 55 (1): 73–86. Bibcode: 2012Palgy..55...73C. doi: 10.1111/j.1475-4983.2011.01117.x.
  53. ^ Clack, Jennifer A. (2011). "A new microsaur from the early carboniferous (Viséan) of East Kirkton, Scotland, showing soft tissue evidence". Special Papers in Palaeontology. 29: 45–55.
  54. ^ Clack, Jennifer A.; Ahlberg, Erik (2004). Arratia, G.; Wilson, M.V.H.; Cloutier, R. (eds.). A new stem tetrapod from the Early Carboniferous of Northern Ireland. München: Verlag Dr. Friedrich Pfeil. pp. 309–320. {{ cite book}}: |work= ignored ( help)
  55. ^ Clack, J A (1 April 2003). "A new baphetid (stem tetrapod) from the Upper Carboniferous of Tyne and Wear, U.K., and the evolution of the tetrapod occiput". Canadian Journal of Earth Sciences. 40 (4): 483–498. Bibcode: 2003CaJES..40..483C. doi: 10.1139/e02-065. ISSN  0008-4077.
  56. ^ Clack, J. A. (1993). "Silvanerpeton miripedes, a new anthracosauroid from the Viséan of East Kirkton, West Lothian, Scotland". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 84 (3–4): 369–376. doi: 10.1017/s0263593300006179. ISSN  1755-6910. S2CID  130790735.
  57. ^ Aucott, Rachel (26 March 2020). "Professor Jenny Clack, FRS, 1947-2020". www.zoo.cam.ac.uk. Department of Zoology, University of Cambridge. Retrieved 27 March 2020.
  58. ^ "Daniel Giraud Elliot Medal". National Academy of Sciences. Archived from the original on 29 December 2010. Retrieved 15 February 2011.
  59. ^ Ahlberg, P. E.; Smithson, T. R. (2021). "Jennifer A. Clack. 3 November 1947—26 March 2020". Biographical Memoirs of Fellows of the Royal Society. 71: 79–101. doi: 10.1098/rsbm.2021.0008. S2CID  234771873.
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  61. ^ "Featured Scientists | Your Inner Fish | PBS". www.pbs.org. Retrieved 30 March 2020.
  62. ^ Koppes, Steve; Allen, Susie (15 May 2013). "University to bestow five honorary degrees at 515th Convocation". U Chicago News. The University of Chicago. Retrieved 3 November 2015.
  63. ^ "Two awards". University of Cambridge - Department of Zoology. 4 July 2013. Retrieved 30 March 2020.
  64. ^ "Figures from public life to be honoured by University of Leicester". University of Leicester. 10 July 2014. Retrieved 3 November 2015.
  65. ^ "Six new members elected to the Academy". 3 February 2014.
  66. ^ "Professor Jenny Clack awarded the Palaeontological Association's Lapworth Medal". University of Cambridge - Department of Zoology. 18 December 2015. Retrieved 28 March 2020.

External links