From Wikipedia, the free encyclopedia
Family of moths
The Heliozelidae , commonly known as shield-bearer moths , are a family of small,
day flying
monotrysian
moths distributed worldwide. The larvae of most heliozelid species are
leaf miners who cut distinctive shield-shaped cases from the surface of the host leaf, hence the common name. Some species are considered pests of commercial crops such as grapevines, cranberries, and walnuts. The taxonomy of this family is poorly understood.
[1]
[2]
Classification
This family includes the following genera and species:
[3]
Antispila
Hübner , 1825
A. ampelopsia Kuroko, 1961
A. argostoma Meyrick, 1916
[4]
A. aristarcha Meyrick, 1916
[4]
A. aurirubra Braun, 1915
A. chlorosema Meyrick, 1931
A. cleyerella Lee, 2006
[5]
A. corniella Kuroko, 1961
A. cornifoliella Clemens, 1860
A. cyclosema Meyrick, 1921
[6]
A. distyliella Lee, Hirowatari & Kuroko, 2006
[7]
A. emeishanensis Liao, Yagi, Hirowatari & Huang, 2019
[8]
A. freemani Lafontaine, 1973
A. grimella (Hübner, 1824)
A. hikosana Kuroko, 1961
A. inouei Kuroko, 1987
A. isabella Clemens, 1860
A. isorrhythma Meyrick, 1926
A. iviella Kuroko, 1961
A. kunyuensis Wang, Liu, Xu & Jiang, 2018
[9]
A. longcangensis Liao, Yagi, Hirowatari & Huang, 2019
[8]
A. merinaella Paulian & Viette, 1955
A. mesogramma Meyrick, 1921
[6]
A. metallella (Denis & Schiffermüller, 1775)
A. nolckeni Zeller, 1877
A. nyssaefoliella Clemens, 1860
A. orbiculella Kuroko, 1961
A. orthodelta Meyrick, 1931
[10]
A. pentalitha Meyrick, 1916
[4]
A. petryi Martini, 1899
A. postscripta Meyrick, 1921
[6]
A. praecincta Meyrick, 1921
[6]
A. purplella Kuroko, 1961
A. sinensis Liu & Wang, 2017
[11]
A. stachjanella Dziurzynski, 1948
A. tateshinensis Kuroko, 1987
A. trypherantis Meyrick, 1916
[4]
A. uenoi Kuroko, 1987
Coptodisca Walsingham, 1895
C. arbutiella Busck, 1904
C. cercocarpella Braun, 1925
C. condaliae Busck, 1900
C. diospyriella (Chambers, 1874)
C. ella (Chambers, 1871)
C. juglandella (Chambers, 1874)
C. juglandiella (Chambers, 1874)
C. kalmiella Dietz, 1921
C. lucifluella (Clemens, 1860)
C. magnella Braun, 1916
C. matheri Lafontaine, 1974
C. negligens Braun, 1920
C. ostryaefoliella (Clemens, 1861)
C. powellella Opler, 1971
C. quercicolella Braun, 1927
C. rhizophorae Walsingham, 1897
C. ribesella Braun, 1925
C. saliciella (Clemens, 1861)
C. splendoriferella (Clemens, 1860)
Heliozela Herrich-Schäffer, 1853
H. aesella (Chambers, 1877)
H. ahenea (Walsingham, 1897)
H. anantia (Meyrick, 1897)
H. anna (Fletcher, 1920)
H. argyrozona (Meyrick, 1918)
H. autogenes Meyrick, 1897
H. castaneella Inoue, 1982
H. catoptrias Meyrick, 1897
H. crypsimetalla Meyrick, 1897
H. cuprea (Walsingham, 1897)
H. eucarpa Meyrick, 1897
H. eugeniella (Busck, 1900)
[13]
H. gracilis (Zeller, 1873)
H. hammoniella Sorhagen, 1885
H. isochroa Meyrick, 1897
H. lithargyrella Zeller, 1850
H. microphylla Meyrick, 1897
H. nephelitis Meyrick, 1897
H. praeustella (Deventer, 1904)
H. prodela Meyrick, 1897
H. resplendella Stainton, 1851
H. rutilella (Walker, 1864)
H. sericiella Haworth
H. siderias Meyrick, 1897
H. sobrinella (Deventer, 1904)
H. subpurpurea (Meyrick, 1934)
H. trisphaera Meyrick, 1897
Hoplophanes
Meyrick , 1897
[15]
H. acrozona Meyrick, 1897
H. aglaodora (Meyrick, 1897)
H. argochalca (Meyrick, 1897)
H. chalcolitha Meyrick, 1897
H. chalcopetala (Meyrick, 1897)
H. chalcophaedra Meyrick, 1897
H. chlorochrysa Meyrick, 1897
H. electritis Meyrick, 1897
H. haplochrysa Meyrick, 1897
H. hemiphragma Meyrick, 1897
H. heterospila Meyrick, 1897
H. memoranda Meyrick, 1897
H. monosema Meyrick, 1897
H. niphochalca Meyrick, 1897
H. panchalca Meyrick, 1897
H. peristera Meyrick, 1897
H. phaeochalca Meyrick, 1897
H. philomacha Meyrick, 1897
H. porphyropla Meyrick, 1897
H. semicuprea Meyrick, 1897
H. tritocosma Meyrick, 1897
References
^ Milla, Liz; van Nieukerken, Erik J.; Vijverberg, Ruben; Doorenweerd, Camiel; Wilcox, Stephen A.; Halsey, Mike; Young, David A.; Jones, Therésa M.; Kallies, Axel; Hilton, Douglas J. (2017).
"A preliminary molecular phylogeny of shield-bearer moths (Lepidoptera: Adeloidea: Heliozelidae) highlights rich undescribed diversity" .
Molecular Phylogenetics and Evolution . 120 .
Academic Press : 129–143.
doi :
10.1016/j.ympev.2017.12.004 .
ISSN
1055-7903 .
^ Milla, Liz; Moussalli, Adnan; Wilcox, Stephen A.; van Nieukerken, Erik J.; Young, David A.; Halsey, Mike; McConville, Thomas; Jones, Therésa M.; Kallies, Axel; Hilton, Douglas J. (2019).
"Phylotranscriptomics resolves phylogeny of the Heliozelidae (Adeloidea: Lepidoptera) and suggests a Late Cretaceous origin in Australia" .
Systematic Entomology . 45 (1).
Royal Entomological Society : 128–143.
doi :
10.1111/syen.12383 .
hdl :
11343/286225 .
ISSN
1365-3113 .
^
"Heliozelidae" .
Global Biodiversity Information Facility . Retrieved 8 December 2023 .
^
a
b
c
d Meyrick, Edward (1916).
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^ Lee, Bong-Woo; Hirowatari, Toshiya; Kuroko, Hiroshi; Arita, Yutaka (2006).
"A new species of the genus Antispila Hubner (Lepidoptera, Heliozelidae), feeding on Cleyera japonica (Theaceae) from Japan" . Lepidoptera Science . 57 (1).
J-STAGE : 7–12.
doi :
10.18984/lepid.57.1_7 .
ISSN
1880-8077 .
^
a
b
c
d
e Meyrick, Edward (1921).
Exotic Microlepidoptera . Vol. 2.
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^ Lee, Bong-Woo; Hirowatari, Toshiya; Kuroko, Hiroshi (2006).
"A New Species of Antispila (Lepidoptera: Heliozelidae) from the Ryukyus, Japan, with Descriptions of Immature Stages" . Annals of the Entomological Society of America . 99 (6).
Oxford University Press : 1019–1027.
doi :
10.1603/0013-8746(2006)99[1019:ANSOAL]2.0.CO;2 .
ISSN
0013-8746 .
^
a
b Liao, Cheng-Qing; Yagi, Sadahisa; Hirowatari, Toshiya; Huang, Guo-Hua (2019). "Two new species of Antispila Hübner (Lepidoptera: Adeloidea: Heliozelidae) associated with Hydrangea (Hydrangeaceae) from China". Journal of Asia-Pacific Entomology . 22 (3).
Elsevier : 941–946.
doi :
10.1016/j.aspen.2019.07.017 .
ISSN
1876-7990 .
^ Wang, Nan; Liu, Tengteng; Xu, Jiasheng; Jiang, Bin (2018).
"The leaf-mining genus Antispila Hübner, 1825 feeding on Vitaceae in Shandong Peninsula, China with one new species (Lepidoptera, Heliozelidae)" .
ZooKeys . 744 .
Pensoft Publishers : 49–65.
doi :
10.3897/zookeys.744.2242 .
ISSN
1313-2970 .
^
a
b
c Meyrick, Edward (1931).
Exotic Microlepidoptera . Vol. 4.
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^ Liu, Tengteng; Wang, Shuxia (2017).
"First report of the leaf-mining genus Antispila Hübner, [1825] from mainland China, with the description of a new species feeding on Cornus (Lepidoptera, Heliozelidae)" .
ZooKeys . 686 .
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doi :
10.3897/zookeys.686.13680 .
ISSN
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PMC
5672564 .
^ van Nieukerken, Erik J.; Wullaert, Steve; Lee, Bong-Woo; Bryner, Rudolf (2021).
"Antispilina ludwigi Hering, 1941 (Lepidoptera, Heliozelidae) a rare but overlooked European leaf miner of Bistorta officinalis (Polygonaceae): new records, redescription, biology and conservation" .
Nota Lepidopterologica . 44 .
Pensoft Publishers : 99–121.
doi :
10.3897/nl.44.63848 .
^
a
b van Nieukerken, Erik J.; Eiseman, Charles S. (2020).
"Splitting the leafmining shield-bearer moth genus Antispila Hübner (Lepidoptera, Heliozelidae): North American species with reduced venation placed in Aspilanta new genus, with a review of heliozelid morphology" .
ZooKeys (957).
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^ van Nieukerken, Erik J.; Geertsema, Henk (2015).
"A new leafminer on grapevine and Rhoicissus (Vitaceae) in South Africa within an expanded generic concept of Holocacista (Insecta, Lepidoptera, Heliozelidae)" .
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^
a
b Meyrick, Edward (1897).
"Descriptions of Australian Microlepidoptera. XVII. Elachistidae" . Proceedings of the Linnean Society of New South Wales . 22 .
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^ Bradley, John David (1961).
"Microlepidoptera from the Solomon Islands. Additional records and descriptions of Microlepidoptera collected in the Solomon Islands by the Rennell Island Expedition 1953-54" .
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