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Genus of gastropods
Helicostyla
A shell of Helicostyla huberorum (holotype at MNHN, Paris)
Scientific classification
Domain:
Eukaryota
Kingdom:
Animalia
Phylum:
Mollusca
Class:
Gastropoda
Subclass:
Heterobranchia
Order:
Stylommatophora
Superfamily:
Helicoidea
Family:
Camaenidae
Subfamily:
Helicostylinae
Genus:
Helicostyla
Férussac , 1821
[1]
Type species
Helix mirabilis Férussac, 1821
Synonyms
Bulimus (Eudoxus) Albers, 1850
Bulimus (Orthostylus) Beck, 1837 (original combination)
Cochlostyla (Helicostyla) Férussac, 1821 (unaccepted combination)
Cochlostyla (Opalliostyla) Pilsbry, 1896 · alternate representation
Cochlostyla (Orthostylus) Beck, 1837 (unaccepted combination)
Eudoxus Albers, 1850
Helicostyla (Cochlodryas) Martens, 1860 · accepted, alternate representation
Helicostyla (Dryocochlias) Möllendorff, 1898 · accepted, alternate representation (basionym)
Helicostyla (Eudoxus) Albers, 1850 · accepted, alternate representation
Helicostyla (Helicostyla) Férussac, 1821 · alternate representation
Helicostyla (Opalliostyla) Pilsbry, 1896 (invalid: unnecessary replacement name)
Helicostyla (Orthostylus) Beck, 1837 · alternate representation
Orthostylus Beck, 1837 (junior synonym)
Helicostyla is a
genus of small, air-breathing
land snails ,
terrestrial
pulmonate
gastropod
mollusks in the subfamily Helicostylinae of the family
Camaenidae . Helicostyla is the
type genus of the subfamily Helicostylinae.
[2]
[3]
Distribution
This genus is
endemic to the
Philippines .
Species
Species within the genus Helicostyla include:
[4]
[5]
Helicostyla albina (Grateloup, 1840)
Helicostyla amagaensis de Chavez, Kendrich, Fontanilla, Batomalaque & Chiba, 2015
Helicostyla amaliae (Möllendorff, 1890 )
Helicostyla aplomorpha (Jonas, 1843)
Helicostyla bicolorata (I. Lea, 1840)
Helicostyla boettgeriana (Möllendorff, 1888)
Helicostyla bullula (Broderip, 1841)
Helicostyla buschi (L. Pfeiffer, 1846)
Helicostyla bustoi (Hidalgo, 1887)
Helicostyla caesar (L. Pfeiffer, 1855)
Helicostyla calamianica (Quadras Moellendorff, 1894)
[6]
Helicostyla canonizadoi (Bartsch, 1932)
Helicostyla chocolatina (Poppe, Tagaro & Sarino, 2015)
Helicostyla cinerosa (L. Pfeiffer, 1855)
Helicostyla collodes (G. B. Sowerby I, 1841)
Helicostyla cunctator (Reeve, 1849)
Helicostyla curta (G. B. Sowerby I, 1841)
Helicostyla daphnis (Broderip, 1841)
Helicostyla decorata (Férussac, 1821)
Helicostyla delicata Fulton, 1903
Helicostyla dimera (Jonas, 1846)
Helicostyla domingoi (Bartsch, 1919)
Helicostyla dimera (Jonas, 1846)
Helicostyla effusa (Pfeiffer, 1842)
Helicostyla euconica (Bartsch, 1932)
Helicostyla fastidiosa (Bartsch, 1932)
Helicostyla faunus (Broderip, 1841)
Helicostyla frater (Férussac, 1821)
Helicostyla fulgens (G. B. Sowerby I, 1841)
Helicostyla fulgetra (Broderip, 1841)
Helicostyla gigas (Hidalgo, 1901)
Helicostyla gilva (Pfeiffer, 1845)
Helicostyla glaucophthalma (L. Pfeiffer, 1851)
Helicostyla halichlora (Semper, 1866)
Helicostyla hololeuca (L. Pfeiffer, 1855)
Helicostyla huberorum Thach, 2021
Helicostyla imperator (Pfeiffer, 1848)
Helicostyla indusiata (L. Pfeiffer, 1871)
Helicostyla jonasi (L. Pfeiffer, 1846)
Helicostyla juglans (L. Pfeiffer, 1842)
Helicostyla lacerata (Semper, 1877) (nomen nudum)
Helicostyla leai (L. Pfeiffer, 1846)
Helicostyla leopardus (L. Pfeiffer, 1845)
Helicostyla leucophaea (G. B. Sowerby I, 1841)
Helicostyla librosa (L. Pfeiffer, 1857)
Helicostyla lignaria (L. Pfeiffer, 1842)
Helicostyla macrostoma (L. Pfeiffer, 1843)
Helicostyla marinduquensis (Hidalgo, 1887)
Helicostyla mateoi (Bartsch, 1932)
Helicostyla mearnsi Bartsch, 1905
Helicostyla mirabilis (Férussac, 1821)
Helicostyla montana (C. Semper, 1877)
Helicostyla mus (Delessert, 1841)
Helicostyla nobilis (Reeve, 1848)
Helicostyla nux (C. Semper, 1877)
Helicostyla okadai Kuroda, 1932
Helicostyla orbitula (Sowerby, 1841)
Helicostyla palavanensis (L. Pfeiffer, 1857)
Helicostyla persimilis (Deshayes, 1850)
Helicostyla philippinensis (L. Pfeiffer, 1846)
Helicostyla pictor (Broderip, 1841)
Helicostyla pithogaster (Ferussac, 1821)
Helicostyla polita Reeve, 1848
Helicostyla portii (L. Pfeiffer, 1861)
Helicostyla rufogaster (Lesson, 1832)
Helicostyla saranganica (Hidalgo, 1887)
Helicostyla satyrus (Broderip, 1841)
Helicostyla schadenbergi (Möllendorff, 1890)
Helicostyla seckendorffiana (L. Pfeiffer, 1847)
Helicostyla simonei Thach & F. Huber, 2021
Helicostyla simplex (Jonas, 1843)
Helicostyla smaragdina (Reeve, 1842)
Helicostyla solida (L. Pfeiffer, 1843)
Helicostyla solivaga (Reeve, 1849)
Helicostyla speciosa (Jay, 1839)
Helicostyla tephrodes (L. Pfeiffer, 1843)
Helicostyla thomsoni (L. Pfeiffer, 1871)
Helicostyla ticaonica (Broderip, 1841)
Helicostyla turbo (Pfeiffer, 1845)
Helicostyla ventricosa (Bruguière, 1792)
Helicostyla vidali (Hidalgo, 1887)
Helicostyla villari (Hidalgo, 1887)
Helicostyla volubilis (Reeve, 1851)
Helicostyla weberi (Bartsch, 1919)
Helicostyla woodiana (I. Lea, 1840)
Helicostyla worcesteri Bartsch, 1909
Species brought into synonymy
Helicostyla aegle (Broderip, 1841) : synonym of
Cochlostyla aegle (Broderip, 1841)
Helicostyla annulata (G.B. Sowerby I, 1841) : synonym of
Pachysphaera annulata (G. B. Sowerby I, 1841) (unaccepted combination)
Helicostyla balteata (G.B. Sowerby I, 1841) : synonym of
Pachysphaera balteata (G. B. Sowerby I, 1841 ) (unaccepted combination)
Helicostyla bembicodes (Pfeiffer, 1851) : synonym of
Helicobulinus bembicodes (L. Pfeiffer, 1851) (unaccepted combination)
Helicostyla butleri (Pfeiffer, 1842) : synonym of
Dryocochlias butleri (L. Pfeiffer, 1842) (unaccepted combination)
Helicostyla camelopardalis (Broderip, 1841) : synonym of
Hypselostyla camelopardalis (Broderip, 1841) (unaccepted combination)
Helicostyla carinata (Lea, 1840) : synonym of
Hypselostyla carinata (I. Lea, 1840) (superseded combination)
Helicostyla cincinna (G.B. Sowerby I, 1840) : synonym of
Hypselostyla cincinna (G. B. Sowerby I, 1841) (unaccepted combination)
Helicostyla cincinniformis (G.B. Sowerb Iy, 1841) : synonym of
Hypselostyla cincinniformis (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla collodes (G.B. Sowerby I, 1841) : synonym of
Cochlostyla collodes (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla concinna (G.B. Sowerby I, 1841) : synonym of
Hypselostyla concinna (G. B. Sowerby I, 1841) (unaccepted combination)
Helicostyla curta (G.B. Sowerby I, 1841) : synonym of
Cochlostyla curta (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla dactylus (Broderip, 1841) : synonym of
Hypselostyla dactylus (Broderip, 1841) (superseded combination)
Helicostyla daphnis (Broderip, 1841) : synonym of
Cochlostyla daphnis (Broderip, 1841) (superseded combination)
Helicostyla decora (A. Adams & Reeve, 1850) : synonym of
Cochlodryas decora (A. Adams & Reeve, 1850) (superseded combination)
Helicostyla denticulata (Jay, 1839) : synonym of
Calocochlea denticulata (Jay, 1839) (superseded combination)
Helicostyla dimera (Jonas, 1846) : synonym of
Cochlostyla dimera (Jonas, 1846) (superseded combination)
Helicostyla diana (Broderip, 1841) : synonym of
Hypselostyla diana (Broderip, 1841) (unaccepted combination)
Helicostyla dubiosa (Pfeiffer, 1846) : synonym of
Cochlostyla dubiosa (L. Pfeiffer, 1846) (superseded combination
Helicostyla euconica (Bartsch, 1932) : synonym of
Cochlostyla effusa (L. Pfeiffer, 1843)
Helicostyla evanescens (Broderip, 1841) : synonym of
Hypselostyla evanescens (Broderip, 1841) (unaccepted combination)
Helicostyla fenestrata (Sowerby I, 1841) : synonym of
Pachysphaera fenestrata (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla florida (Sowerby I, 1841) : synonym of
Cochlodryas florida (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla fulgens (Broderip, 1841) : synonym of
Cochlostyla fulgens (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla fulgetra (Broderip, 1841) : synonym of
Cochlostyla fulgetra (Broderip, 1841)
Helicostyla grandis (Pfeiffer, 1845) : synonym of
Helicobulinus grandis (L. Pfeiffer, 1845) (unaccepted combination)
Helicostyla hydrophana G.B. Sowerby I, 1841 : synonym of
Dryocochlias metaformis (Férussac, 1821) (junior synonym)
Helicostyla ignobilis Sowerby I, 1841 : synonym of
Cochlodryas ignobilis (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla iloconensis (Sowerby I, 1841) : synonym of
Pachysphaera iloconensis (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
Helicostyla jonasi (Pfeiffer, 1845) : synonym of
Cochlostyla jonasi (L. Pfeiffer, 1846) (superseded combination)
Helicostyla juglans (Pfeiffer, 1842) : synonym of
Cochlostyla juglans (L. Pfeiffer, 1842) (superseded combination)
Helicostyla leopardus (Pfeiffer, 1845) : synonym of
Cochlostyla leopardus (L. Pfeiffer, 1845) (superseded combination)
Helicostyla leucophaea (Sowerby I, 1841) : synonym of
Cochlostyla leucophaea (G. B. Sowerby I, 1841) (superseded combination)
Helicostyla lignaria (Pfeiffer, 1842) : synonym of
Cochlostyla lignaria (L. Pfeiffer, 1842) (suspected synonym)
Helicostyla macrostoma (Pfeiffer, 1843) : synonym of
Cochlostyla macrostoma (L. Pfeiffer, 1843) (superseded combination)
Helicostyla marinduquensis (Hidalgo, 1887) : synonym of
Cochlostyla marinduquensis Hidalgo, 1887 (superseded combination)
Helicostyla metaformis (Ferussac, 1821) : synonym of
Dryocochlias metaformis (Férussac, 1821) (misspelling of original name, Corasia, Albers, 1850)
Helicostyla mindoroensis W. J. Broderip, 1892 : synonym of
Chrysallis mindoroensis (Broderip, 1841) (unaccepted generic combination)
Helicostyla mirabilis (Ferussac, 1821) : synonym of
Cochlostyla mirabilis (Férussac, 1821) (superseded combination)
Helicostyla montana Semper, 1891 : synonym of
Cochlostyla montana C. Semper, 1877 (superseded combination)
Helicostyla monticula (G.B. Sowerby I, 1841) : synonym of
Orustia monticula (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
Helicostyla nimbosa (Broderip, 1841) : synonym of
Hypselostyla nimbosa (Broderip, 1841) (unaccepted combination)
Helicostyla olanivanensis (Bartsch, 1913) : synonym of
Cochlostyla olanivanensis Bartsch, 1913
Helicostyla opalina (G.B. Sowerby I, 1841) : synonym of
Phengus opalinus (G. B. Sowerby I, 1841)
Helicostyla phaeostyla (Pfeiffer, 1856) : synonym of
Cochlostyla phaeostyla (L. Pfeiffer, 1857)
Helicostyla pictor (Broderip, 1841) : synonym of
Cochlostyla pictor (Broderip, 1841) (superseded combination)
Helicostyla polillensis (Pfeiffer, 1861) : synonym of Cochlostyla polillensis (L. Pfeiffer, 1861) accepted as
Calocochlea polillensis (L. Pfeiffer, 1861)
Helicostyla pudibunda Semper, 1891 : synonym of
Pfeifferia pudibunda (C. Semper, 1877) (unaccepted combination)
Helicostyla rehbeini (L. Pfeiffer, 1852) : synonym of
Dryocochlias rehbeini (L. Pfeiffer, 1852) (unaccepted combination)
Helicostyla rufogaster (Lesson, 1831) : synonym of
Cochlostyla rufogaster (Lesson, 1832) (superseded combination)
Helicostyla rustica (Mousson, 1849) : synonym of Dryocochlias metaformis rustica (Mousson, 1849)
Helicostyla sarcinosa (Ferussac, 1821) : synonym of
Helicobulinus sarcinosa (Férussac, 1821)
Helicostyla satyra (Broderip, 1841) : synonym of
Cochlostyla satyrus (Broderip, 1841) (superseded combination)
Helicostyla simplex (Jonas, 1843) : synonym of
Cochlostyla simplex (Jonas, 1843) (superseded combination)
Helicostyla solivaga (Reeve, 1849) : synonym of
Cochlostyla solivaga (Reeve, 1849) (superseded combination)
Helicostyla sphaerica (Sowerby I, 1841) : synonym of
Pachysphaera sphaerica (G. B. Sowerby I, 1841)
Helicostyla submirabilis Möllendorff, 1897 : synonym of
Cochlostyla submirabilis Möllendorff, 1897 (unaccepted combination)
* Helicostyla turbinoides (Broderip, 1841) : synonym of
Helicobulinus turbinoides (Broderip, 1841) (superseded combination)
Helicostyla velata (Broderip, 1841) : synonym of
Hypselostyla velata (W.J. Broderip, 1841)
Helicostyla ventricosa (Bruguière, 1792) : synonym of
Cochlostyla ventricosa (Bruguière, 1792) (superseded combination)
Helicostyla viridostriata (Lea, 1840) : synonym of
Cochlodryas viridostriata (I. Lea, 1840) (superseded combination)
Helicostyla woodiana (Lea, 1840) : synonym of
Cochlostyla woodiana (I. Lea, 1840) (superseded combination)
References
^ Férussac A. É. d'A. de (1821). Histoire naturelle générale et particulière des mollusques terrestres et fluviatiles , Tabl. Limaçons, 28.
^
Bouchet, Philippe ;
Rocroi, Jean-Pierre ; Frýda, Jiri; Hausdorf, Bernard;
Ponder, Winston ; Valdés, Ángel & Warén, Anders (2005). "
Classification and nomenclator of gastropod families ".
Malacologia . 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397.
ISBN
3-925919-72-4 .
ISSN
0076-2997 .
^ MolluscaBase eds. (2021). MolluscaBase. Helicostyla Férussac, 1821. Accessed through: World Register of Marine Species at:
http://www.marinespecies.org/aphia.php?p=taxdetails&id=993127 on 2021-02-02
^
Bradybaenidae . Malacolos.com, accessed 16 November 2010.
^
Biolib
^ Springsteen, F.J. & Leobrera, F.M. 1986. Shells of the Philippines. Carfel Seashell Museum, Philippines: 1-377.
Bank, R. A. (2017). Classification of the Recent terrestrial Gastropoda of the World. Last update: July 16, 2017
External links
Media related to
Helicostyla at Wikimedia Commons
Férussac, A.E.J.P.F. d'Audebard de. (1821-1822). Tableaux systématiques des animaux mollusques classés en familles naturelles, dans lesquels on a établi la concordance de tous les systèmes; suivis d'un Prodrome général pour tous les mollusques ou fluviatiles, vivantes ou fossiles. Paris, 1821 et 1822.
Pilsbry, H. A. (1896). On the names of certain subgenera of Helicostyla. The Nautilus. 9(9): 108.
Albers, J. C. (1850). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet. Berlin: Enslin. 262 pp
Kennard, A. S. (1942). The Histoire and Prodrome of Férussac. Part III. The divisional names. Proceedings of the Malacological Society of London. 25 (3): 111-118. London
Pilsbry, H. A. (1903). The use of the generic name Helicostyla. The Nautilus. 17(5): 58
Albers, J. C.; Martens, E. von. (1860). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet von Joh. Christ. Albers. Ed. 2. Pp. i-xviii, 1-359. Leipzig: Engelman